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The Predaceous Diving Beetles (Coleoptera: Dytiscidae) of Papua Indonesia
by Michael Balke
Introduction
Dytiscidae are the predaceous diving beetles, a family of water beetles. They are about 25 mm long on average, though there is much variation between species. Most are dark brown, blackish or dark olive in color with golden highlights in some subfamilies. They have short, but sharp mandibles. Immediately upon biting they deliver digestive enzymes. The larvae are commonly known as water tigers. The family has not been comprehensively cataloged since 1920, but is estimated to include about 4,000 species in over 160 genera.
Larvae vary in size from about 1 to 5 cm. The larvae's bodies are shaped like crescents, with the tail long and covered with thin hairs. Six legs protrude from along the belly, which also sports the same thin hairs. The head is flat and square, with a pair of long, large pincers. When hunting, they cling to grasses or pieces of wood along the bottom, and hold perfectly still until prey passes by, then they lunge, trapping their soon-to-be-food between their front legs and biting down with its pincers. Their usual prey includes tadpoles and glassworms, among dozens of other smaller water-dwelling creatures.
As soon as the beetles come to the stage in life in which they mature to adulthood, the larva crawl from the water on the sturdy legs, and bury themselves in the mud for pupation. After about a week, or longer in some species, they emerge from the mud as adults. Large but slow on land and not particularly fierce as adults, they are also eaten with relish by many mid-sized birds, mammals and other larger predators. The larvae are usually safer, due to their camouflage and ability to escape by water jet; they can be quite hard to catch and may become apex predators in small ponds.
see Wikipedia for more information
Listed Papua species (with certainty): 48 [species endemic for Papua are not defined yet]
grey coloured taxa are found in Papua New Guinea or known from "New Guinea" but are not confirmed from Papua Indonesia yet
Copelatinae
Tribus Copelatini
Copelatus Erichson, 1832
Sg. Copelatus Erichson, 1832biroi Guignot, 1956
gentilis Sharp, 1882
gestroi Régimbart, 1892
neoguinensis Zimmermann, 1919
irregularis MacLeay, 1825
kaszabi Guignot, 1956
luteomaculatus Guignot, 1956
papuensis Balfour-Browne, 1939
portior Guignot, 1956
tulagicus Guignot, 1942
apicalis Balfour-Browne, 1939vagestriatus Zimmermann, 1919
xanthocephalus Régimbart, 1899
Exocelina Balke, 1998
aipo Balke, 1998
aipomek Balke, 1998
ascendens Balke, 1998
astrophallus Balke, 1998
atratus Balfour-Browne, 1939
casuarinus Balke, 1998
bacchusi Balke, 1998
broschii Balke, 1998
damantiensis Balke, 1998
danae Balke, 1998
desii Balke, 1999
erteldi Balke, 1998
formosus Balke, 1998
fume Balke, 1998
heidiae Balke, 1998
inornatus Balke, 1998
jaseminae Balke, 1998
karmurensis Balke, 1998
ketembang Balke, 1998
larsoni Balke, 1998
manfredi Balke, 1998
me Balke, 1998
messeri Balke, 1999
miriae Balke, 1998
monae Balke, 1998
nomax Balfour-Browne, 1939
patepensis Balke, 1998
politus Sharp, 1882
rivulus Balke, 1998
rufus Balke, 1998
sanctimontis Balke, 1998
speciosus Balke, 1998
takime Balke, 1998
talaki Balke, 1998
tarmluensis Balke, 1998
ullrichi Balke, 1998
HydroporinaeTribus Hyphydrini
Hyphydrus Illiger, 1802
bistrimaculatus Biström & Biström, 1997
dani Biström, Balke & Hendrich, 1993
decemmaculatus Wehncke, 1877
curvipes Régimbart, 1892elegans (Montrouzier, 1860)
gibbicollis Régimbart, 1892gibbosus Biström, 1984
loriae Régimbart, 1892
lyratus Swartz, 1808
Tribus Hydrovatini
Hydrovatus Motschulsky, 1853
enigmaticus Biström, 1996
fasciatus Sharp, 1882
fractus Sharp, 1882
laosensis Biström, 1996
opacus Sharp, 1882
nigrita Sharp, 1882
irianensis Biström, 1996
samuelsoni Biström, 1996
Tribus Bidessini
Clypeodytes Régimbart, 1894
loriae (Régimbart, 1892)
Hydroglyphus Motschulsky, 1853baeri (Régimbart, 1895)
Limbodessus Guignot, 1939compactus (Clark, 1862)
neoguinensis Régimbart, 1892Tribus Hydroporini
Chostonectes Sharp, 1882
maai Balke, 1995
Megaporus Brinck, 1943piceatus (Régimbart, 1892)
Sternopriscus Sharp, 1882hansardi (Clark, 1862)
LaccophilinaeTribus Laccophilini
Laccophilus Leach, 1815
bapak Balke, Larson & Hendrich, 1996
cingulatus Sharp, 1882
clarki Sharp, 1882
ekari Balke, Larson & Hendrich, 1996
heidiae Brancucci, 1983
papuanus Balke, Larson & Hendrich, 1996
quadrimaculatus Sharp, 1882 sensu lato
ramuensis Balke, Larson & Hendrich, 1996
schereri Brancucci, 1983
seminiger Fauvel, 1883
auriculatus Régimbart, 1892transversalis Régimbart, 1877
univittatus Régimbart, 1891
ameliae Balke et al., 2000
ssp. ameliae Balke et al., 2000
ssp. weylandensis Balke et al., 2000
aterrimus Balke et al., 2000bacchusi Balke et al., 2000
bellissimus Balke et al., 2000
bicinctus (Régimbart, 1892)
irianensisi Balke et al., 2000
kokodanus Balke et al., 2000
ramuensis Balke et al., 2000
speciosus (Régimbart, 1892)
ColymbetinaeTribus Agabini
Platynectes Régimbart, 1878beroni Guéorguiev, 1978
decastigma Régimbart, 1899
decempunctatus (Fabricius, 1775)
octodemimmaculatus (MacLeay, 1825)
neoguinensis Guéorguiev & Rocchi, 1993
semperi Régimbart, 1899
Tribus Colymbetini
Carabdytes Balke, Hendrich & Wewalka, 1992
upin Balke, Hendrich & Wewalka, 1992Rhantus Dejean, 1833
anggi Balke 2001
bacchusi Balke et al., 2000
dani Balke et al., 2000
ekari Balke & Hendrich, 1992
kakapupu Balke et al., 2000
papuanus Balfour-Browne, 1939
riedeli Balke et al., 2000
supranubicus Balke et al., 2000
suturalis (MacLeay, 1825)
neoguinensis Guéorguiev & Rocchi, 1993DytiscinaeTribus Eretini
Eretes Laporte, 1833
sticticus (L., 1767)Tribus Hydaticini
Hydaticus Leach, 1817
okalehubyi Balke & Hendrich, 1992
quadrivittaus Blanchard, 1853
rhantaticoides Régimbart, 1892
rivanolis Wewalka, 1979
sellatus Régimbart, 1883
suffusus Régimbart, 1892
Tribus Acilini
Rhantaticus Sharp, 1882
congestus (Klug, 1832)Sandracottus Sharp, 1882
guerini Balfour-Browne, 1939
flavocinctus Guérin,1836, nom. praeocc.
femoralis Heller, 1934 nom. praeocc.Tribus Cybistrini
Homoeodytes Régimbart, 1878
atratus (Fabricius, 1801)
gibbosus Wilke, 1919
Cybister Curtis, 1827
godeffroyi Wehncke, 1876
hypomelas Régimbart, 1892
kolbei Wilke, 1919 n.comb.
loxidiscus Wilke, 1919
papuanus Guignot, 1956
tripunctatus (Olivier, 1795)
sugillatus Erichson, 1834
yulensis Guignot, 1956
last updated on 24th December 2010
Literature
Balke M. 2000. Die Schwimmkäfer Neu Guineas. Artenreichtum, Phylogenie, Biogeographie und Lebensweise (Coleoptera: Dytiscidae). Dissertation.de, Berlin, Germany.