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The Papua Insects Foundation
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The Scale Insects (Hemiptera: Sternorrhyncha, Coccinea) of Papua Indonesia
by Ilya A. Gavrilov-Zimin
Listed Papua species: 71; species in grey have only been found in Papua New Guinea.
ORTHEZIIDAENeonipponorthezia Kozár, 2004
kaindii Konczné Benedicty et Kozár, 2004
regina Konczné Benedicty et Kozár, 2004
Newsteadia Green, 1902
biroi Kozár et Konczné Benedicty, 2000
guineensis Kozár et Konczné Benedicty, 2000
milleri Kozár et Konczné Benedicty, 2000
Nipponorthezinella Kozár, 2004
guadalcanalia Morrison, 1952
MARGARODIDAE s.l.
Monophlebinae
Etropera Bhatti et Gullan, 1990
ablusa Bhatti et Gullan, 1990
papuensis Bhatti et Gullan, 1990 [endemic]
pilosa Bhatti et Gullan, 1990
sedlaceki Bhatti et Gullan, 1990
Icerya Signoret, 1876
gaponiana Gavrilov-Zimin, 2013 [endemic]
purchasi Maskell, 1879
imperatae Rao, 1951
samaraia Morrison, 1927
seychellarum Westwood, 1855
Modicicoccus Bhatti et Gullan, 1990
gagnei Bhatti et Gullan, 1990
kaindiensis Bhatti et Gullan, 1990
monticolus Bhatti et Gullan, 1990
newsteadi Bhatti et Gullan, 1990
Monophlebulus Cockerell, 1902
enarotalicus Bhatti et Gullan, 1990 [endemic]
gressitti Bhatti et Gullan, 1990 [endemic]
montanus Reyne, 1965
toxopei Reyne, 1965
Peengea Bhatti et Gullan, 1990
affinis Bhatti et Gullan, 1990
Margarodinae
Margarodes Guilding, 1829
australis (Jakubski, 1965)
PSEUDOCOCCIDAE
Phenacoccinae
Brevennia Goux, 1940
rehi (Lindinger, 1943)
Laingiococcus Morrison, 1945
painei (Laing, 1930)
Phenacoccus Cockerell, 1893
solenopsis Tinsley, 1898
Rastrococcus Ferris, 1954
biggeri Williams et Watson, 1988
expeditionis Williams, 1989
neoguineensis Williams et Watson, 1988 [endemic]
vicorum Williams et Watson, 1988
Pseudococcinae
Antonina Signoret, 1875
graminis Maskell, 1897
pretiosa Ferris, 1953
Cannococcus Borchsenius, 1960
ikshu Williams et Watson, 1988
Chaetococcus Maskell, 1898
bambusae Maskell, 1893
Chorizococcus McKenzie, 1960
talipikanus Williams et Watson, 1988
Criniticoccus Williams, 1960
theobromae Williams, 1960
Crisicoccus Ferris, 1950
chiponensis Takahashi, 1939
strigilis Gavrilov-Zimin, 2013 [endemic]
theobromae Williams et Watson, 1988
Dysmicoccus Ferris, 1950
amnicola Williams et Watson, 1988
brevipes (Cockerell, 1893)
busoensis Williams et Watson, 1988
comitatus Williams et Watson, 1988 [endemic]
hylonomus Williams et Watson, 1988
maclayi Gavrilov-Zimin, 2013 [endemic]
nesophilus Williams et Watson, 1988
papuanicus Williams et Watson, 1988
Ferrisia Fullaway, 1923
malvastra (McDaniel, 1962)
virgata (Cockerell, 1893)
Leptococcus Reyne, 1961
metroxyli Reyne, 1961 [endemic]
Maconellicoccus Ezzat, 1958
hirsutus Green, 1908
Maculicoccus Williams, 1960
malaitensis (Cockerell, 1929)
Madangiacoccus Williams, 1985
araucariae Williams, 1985
Mollicoccus Williams, 1960
guadalcanalanus Williams, 1960
Mutabilicoccus Williams, 1960
simmondsi (Laing, 1925)
vanheurni (Reyne, 1961) [endemic]
Nipaecoccus Šulc, 1945
viridis (Newstead, 1894)
Palmicultor Williams, 1963
browni (Williams, 1960)
lumpurensis Takahashi, 1951
Papuacoccus Williams et Watson, 1988
barymelus Williams et Watson, 1988
Paracoccus Ezzat et McConnell, 1956
interceptus Lit, 1997
melanesicus Williams et Watson, 1988
morobensis Williams et Watson, 1988
Paraputo Laing, 1929
chimbuensis Williams, 2005 [endemic]
leveri (Green, 1934)
Planococcus Ferris, 1950
citri (Risso, 1813)
dioscoreae Williams, 1960
lilacinus (Cockerell, 1905)
minor (Maskell, 1897)
Pseudococcus Westwood, 1840
araucariarum Williams, 1985
daymananus Williams, 1985
dendrobiorum Williams, 1985
grayi Williams, 1985 [endemic]
jackbeardsleyi Gimpel et Miller, 1996
linearis Williams et Watson, 1988
longispinus (Targioni Tozzetti, 1867)
queenslandicus Williams, 1985
saccharicola Takahashi, 1928
solomonensis Williams, 1960
Saccharicoccus Ferris, 1950
sacchari (Cockerell, 1895)
Trionymus Berg, 1899
palauensis Beardsley, 1966
Rhizoecinae
Geococcus Green, 1902
baloghi Kianek et Konczné Benedicty, 2007
coffeae Green, 1933
Ripersiella Tinsley, 1899
carolinensis (Beardsley, 1966)
guineensis Kozár et Konczné Benedicty, 2003
puhiensis (Hambleton, 1974)
Xenococcus Silvestri, 1924
acropygae Williams, 1998
annandalei Silvestri, 1924
neoguineensis (Williams), 1998
ERIOCOCCIDAE
Acanthococcus Signoret, 1875
araucariae (Maskell, 1879)
Apiomorpha Rübsaamen, 1894
pedunculata (Fuller, 1896)
Opisthoscelis Schrader, 1863
subrotunda Schrader, 1863
Pseudomontanococcus Kozár et al., 2008
baloghi Kozár et al., 2008 [endemic]
martini Kozár et al., 2008 [endemic]
Sangicoccus Reyne, 1961
obtusispinus (Reyne, 1961) [endemic]
truncatispinus (Reyne, 1961)
COCCIDAE
Anthococcus Williams et Watson, 1990
keravatae Williams et Watson, 1990
Ceroplastes Gray, 1828
ceriferus (Fabricius, 1798)
destructor Newstead, 1917
floridensis Comstock, 1881
murrayi Froggatt, 1919
rubens Maskell, 1893
rusci (Linnaeus, 1758)
stellifer (Westwood, 1871)
Coccus Linnaeus, 1758
celatus De Lotto, 1960
hesperidum Linnaeus, 1758
longulus (Douglas, 1887)
viridis (Green, 1889)
Drepanococcus Williams et Watson, 1990
chiton (Green, 1908)
Eucalymnatus Cockerell, 1901
tessellatus (Signoret, 1873)
Fistulococcus Hodgson et Martin, 2005
intsiae Hodgson et Martin
Kilifia De Lotto, 1965
acuminata (Signoret, 1873)
Luzulaspis Cockerell, 1902
australis (Maskell, 1894)
Megapulvinaria Yang, 1982
maxima (Green, 1904)
Melanesicoccus Williams et Watson, 1990
kleinhoviae Williams et Watson, 1990
Milviscutulus Williams et Watson, 1990
ciliatus Williams et Watson, 1990
mangiferae (Green, 1889)
pilosus Williams et Watson, 1990
spiculatus Williams et Watson, 1990
Myzolecanium Beccari, 1877
kibarae Beccari, 1877
robertsi (Williams et Watson, 1990)
Neosaissetia Tao et al., 1983
keravatae Williams et Watson, 1990
Paralecanium Cockerell, 1899
ovatum Morrison, 1921
Parasaissetia Takahashi, 1955
nigra (Nietner, 1861)
Platylecanium Cockerell & Robinson, 1915
cocotis Laing, 1925
Prococcus Avasthi, 1993
acutissimus (Green, 1896)
Pulvinaria Targioni Tozzetti, 1866
cacao Williams et Watson, 1990 [endemic]
elongata Newstead, 1917
psidii Maskell, 1893
urbicola Cockerell, 1893
Saissetia Deplanche, 1859
coffeae (Walker, 1852)
miranda (Cockerell, 1899)
vivipara Williams et Watson, 1990
KERRIIDAEParatachardina Balachowsky, 1950
morobensis Williams et Watson, 1990
LECANODIASPIDIDAE
Lecanodiaspis Targioni Tozzetti, 1869
casuarinae Williams et Watson, 1990
Psoraleococcus Borchsenius, 1959
browni Lambdin et Kosztarab, 1988
ASTEROLECANIIDAE
Asterolecanium Targioni Tozzetti, 1868
neoguineense Williams et Watson, 1990 [endemic]
Bambusaspis Cockerell, 1902
bambusae (Boisduval, 1869)
Pauroaspis Tang, 1995
daedalea Gavrilov-Zimin, 2013 [endemic]
Russellaspis Bodenheimer, 1951
pustulans (Cockerell, 1892)
ACLERDIDAE
Aclerda Signoret, 1874
pseudozoysiae Gavrilov-Zimin, 2012 [endemic]
DIASPIDIDAE
Abgrallaspis Balachowsky, 1948
cyanophylli (Signoret, 1869)
ruebsaameni (Cockerell, 1902)
Allantomytilus Leonardi, 1898
dacryoides Williams et Watson, 1988 [endemic]
Andaspis MacGillivray, 1921
maai Williams et Watson, 1988
numerata Brimblecombe, 1959
spinosa Williams et Watson, 1988
Aonidiella Berlese et Leonardi, 1896
aurantii (Maskell, 1879)
citrina (Coquillett, 1891)
comperei McKenzie, 1937
eremocitri McKenzie, 1937
inornata McKenzie, 1938
orientalis (Newstead, 1894)
Aspidiella Leonardi, 1898
hartii (Cockerell, 1895)
sacchari (Cockerell, 1893)
Aspidiotus Bouché, 1833
destructor Signoret, 1869
excisus Green, 1896
musae Williams et Watson, 1988
Aulacaspis Cockerell, 1893
australis Brimblecombe, 1959
madiunensis (Zehntner, 1898)
neoguineensis Williams et Watson, 1988
rosarum Borchsenius, 1958
tegalensis (Zehntner, 1898)
Chionaspis Signoret, 1868
broughae Williams et Watson, 1988
comys Williams et Watson, 1988 [endemic]
keravatana Williams et Watson, 1988
Chrysomphalus Ashmead, 1880
aonidum (Linnaeus, 1758)
dictyospermi (Morgan, 1889)
pinnulifer (Maskell, 1891)
Diaspis Costa, 1828
echinocacti (Bouché, 1791)
Duplaspidiotus MacGillivray, 1921
claviger (Cockerell, 1901)
Fiorinia Targioni Tozzetti, 1868
biakana Williams et Watson, 1988 [endemic]
coronata Williams et Watson, 1988
reducta Williams et Watson, 1988 [endemic]
theae Green, 1900
Furcaspis Lindinger, 1908
exophthalma Williams et Miller, 2006
Hemiberlesia Cockerell, 1897
lataniae (Signoret, 1869)
palmae (Cockerell, 1892)
Howardia Berlese et Leonardi, 1896
biclavis (Comstock, 1883)
Ischnaspis Douglas, 1887
longirostris (Signoret, 1882)
Lepidosaphes Shimer, 1868
beckii (Newman, 1869)
elmerrilleae Williams et Watson, 1988
eurychlidonis Williams et Watson, 1988
gloverii (Packard, 1869)
karkarica Williams et Watson, 1988
rubrovittata Cockerell, 1905
securicula Williams et Watson, 1988
tapleyi Williams, 1960
tokionis (Kuwana, 1902)
Lopholeucaspis Balachowsky, 1953
baluanensis Williams et Watson, 1988
Morganella Cockerell, 1897
longispina (Morgan, 1889)
Octaspidiotus MacGillivray, 1921
australiensis (Kuwana, 1931)
Odonaspis Leonardi, 1897
ruthae Kotinsky, 1915
saccharicaulis (Zehntner, 1897)
Parlatoria Targioni Tozzetti, 1868
araucariae (Williams et Watson, 1988)
crotonis Douglas, 1887
proteus (Curtis, 1843)
pseudaspidiotus Lindinger 1905
ziziphi (Lucas, 1853)
Pinnaspis Cockerell, 1892
aspidistrae (Signoret, 1869)
buxi (Bouché, 1851)
strachani (Cooley, 1899)
Pseudaulacaspis MacGillivray, 1921
cockerelli (Cooley, 1897)
multiducta Williams et Watson, 1988 [endemic]
papulosa Williams et Watson, 1988 [endemic]
pentagona (Targioni Tozzetti, 1886)
ponticula Williams et Watson, 1988
Rugpapuaspis Ben-Dov, 1991
proxantennata Ben-Dov, 1991
Schizentaspidus Mamet, 1958
silvicola Williams et Watson, 1988
Silvestraspis Bellio, 1929
ficaria Williams et Watson, 1988
uberifera Lindinger, 1911
Unaspis MacGillivray, 1921
citri (Comstock, 1883)
Checklist updated on 8th September 2013
Literature
Ben-Dov, Y., Miller, D.R., Gibson, G.A.P., 2012. ScaleNet. A database of the scale insects of the world. Available at: http://www.sel.barc.usda.gov/scalenet/scalenet.htm (last updated 28 August 2013).
Bhatti, S. & Gullan, P.J., 1990. New margarodid species (Homoptera: Coccoidea: Margarodidae: Monophlebinae) from New Guinea. Invertebrate Taxonomy 3:877-911.
Gavrilov-Zimin, I.A., 2012. A contribution to the taxonomy, cytogenetics and reproductive biology of the genus Aclerda Signoret (Homoptera, Coccinea, Aclerdidae). Comparative Cytogenetics 6(4): 389-395.
Gavrilov-Zimin, I.A., 2013. New scale insects (Homoptera: Coccinea) from Sulawesi and New Guinea, with some other additions to the Indonesian fauna. Tropical Zoology 26(2):64-86.
Williams, D.J., 2004. Mealybugs of southern Asia. Kuala Lumpur: Southdene SDN.
Williams, D.J. & Watson, G.W., 1988a. The Scale Insects of the Tropical South Pacific Region. Pt. 1. The Armoured Scales (Diaspididae). Wallingford, UK: CAB International.
Williams, D.J. & Watson, G.W., 1988b. The Scale Insects of the Tropical South Pacific Region. Pt. 2: The Mealybugs (Pseudococcidae). Wallingford, UK: CAB International.
Williams, D.J. & Watson, G.W., 1990. The Scale Insects of the Tropical South Pacific Region. Pt. 3: The Soft Scales (Coccidae) and Other Families. Wallingford, UK: CAB International.
