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The Ground Beetles (Coleoptera: Carabidae) of Papua Indonesia
by Martin Baehr
IntroductionThe present list of species recorded from West Papua is principally based on P. J. Darlington’s monumental work “The Carabid Beetles of New Guinea” which was printed in four parts between 1952 and 1971, and on the material which I identified, or mentioned and described in a number of descriptive and revisionary papers. It only includes printed records. Apart from that, certainly the list is quite incomplete for a number of other reasons:
1. Only a part of the material of West Papuan Carabidae present in collections has been examined by me and probably not all published records are noted herein. Examination of specimens stored in a number of large museum collections (Natural History Museum, Museum National de l’Histoire Naturelle, B. P. Bishop Museum, Museum of Comparative Zoology, and others) certainly would raise the number of species recorded. At present, however, this is outside of the scope of this list.
2. Even with the recent exploration of several areas within West Papua by a number of German, Dutch, Italian, American and other collectors, knowledge of the carabid fauna of New Guinea in general, and of that of West Papua in particular, still is very unsatisfactory and certainly some species recorded so far only from the eastern half of New Guinea (Papua New Guinea) in future will be detected in the western half (West Papua) and vice versa, which means that the number of already described species occurring in West Papua most probably will raise, but the number of endemic ones among these will decrease.
3. In general, the carabid fauna of West Papua still is much less represented in collections and therefore even less known than that of Papua New Guinea, and moreover, the representation of tribes in collections and in the literature is rather unbalanced. This is due to the still much poorer exploration of West Papua as compared with Papua New Guinea which was sampled by many more collectors not only in early times, but also during the second half of last Century.
4. Although the exploration of West Papua in recent times was much improved through expeditions conducted by a number of keen collectors, large areas in both parts of New Guinea have been left almost untouched and most collectors are visiting well known, rather easily accessible, and relatively safe regions.
5. Almost all recent material which I have examined was sampled by non-carabidologists, namely collectors of Lepidoptera and of diverse groups of non-carabid Coleoptera (weevils, longicorn beetles, water beetles, and else). Therefore sampling methods suitable for carabid beetles were rarely conducted, or these collectors brought in rather carabid beetles of certain life style types. Sampling by use of pitfall traps, digging in the ground, collecting in sand and gravel near water, fogging canopy or tree trunks, and collecting at blacklight was little or not conducted and therefore certain carabid groups are much underrepresented in the samples of the mentioned collectors.
6. Recent revisions of certain genera reveal the existence of very large numbers of species in addition to those mentioned or described by Darlington. Hence already the complete revision of the material existing in collections certainly would multiply the number of species in many genera. Dissections of male genitalia in material of some of the larger genera available to me and conducted by me recently (e.g. Clivina, Tachys, Prosopogmus, Altagonum, Maculagonum, Notagonum, Catascopus, Demetrida) reveal a multitude of externally very similar species that exhibit striking differences in their genitalia and hence represent many additional species.Conclusion
Although Darlington’s monumental revision covers almost 700 carabid species for whole New Guinea, I estimate the actually existing number at least twice but perhaps even three times as large. Hence the number of species occurring in West Papua most probably is well over 1.000, whereas the list below at present covers 532 taxa (species and subspecies) as recorded from West Papua.
For a quick search to subfamily click on: Paussinae, Scaritinae, Trechinae, Harpalinae, Oodinae, Cyclosominae, Brachininae
Listed Papua species: 617 [262 species endemic for Papua]
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Tribus Ozaenini
Pseudozaena Castelnau, 1834
Tribus Scaritini
Clivina Latreille, 1802
alternans Darlington, 1971
biroi Kult, 1951
cardiothorax Baehr, 2008
csikii Kult, 1951
deleta Darlington, 1962
dealata Darlington, 1962
ssp. brachyptera Darlington, 1962 [endemic]erugata Darlington, 1962
komareki Kult, 1951
kulti Darlington, 1962
puncticeps Darlington, 1962 [endemic]
sansapor Darlington, 1962 [endemic]
subfusa Darlington, 1962
szekessyi Kult, 1951
toxopei Darlington, 1962 [endemic]
tripuncta Darlington, 1962
Geoscaptus Chaudoir, 1855cacus (Macleay, 1863)Syleter Andrews, 1941papua Darlington, 1962Tribus Bembidiini
Desarmatocillenus Netolitzky, 1939
albertisi (Putzeys, 1875)Tachys s.l. Stephens, 1828apex Darlington, 1962
aeneus Putzeys, 1875
beatus Darlington, 1962
ceylanicus (Nietner, 1858)
cruciger Putzeys, 1875
curticollis Soane, 1896
erotyloides Andrewes, 1925
fasciatus (Motschulsky, 1851)
fumatus Darlington, 1962
fumax Darlington, 1962
fumicatus Motschulsky, 1851
latissimus Motschulsky, 1851
ssp. tinctus Darlington, 1962 [endemic]loriae Andrewes, 1925
luscus Darlington, 1962 [endemic]
masculus Darlington, 1962
ssp. masculus Darlington, 1962
ssp. filius Darlington, 1962 [endemic]mastersii Macleay, 1871
ssp. exul Darlington, 1962 [endemic]
ssp. pinguis Darlington, 1962nitens Andrewes, 1925
par Darlington, 1962 [endemic]
parapictus Darlington, 1962
parasenarius Darlington, 1971 [endemic]
pictus Andrewes, 1925
ssp. pictus Andrewes, 1925
ssp. subpictus Darlington, 1962 [endemic]plagiatus Putzeys, 1875
psiloides Darlington, 1962
quadrillum Schaum, 1860
serra Darlington, 1962
ssp. breviserra Darlington, 1962 [endemic]
ssp. latiserra Darlington, 1962subbrunneus Darlington, 1962
sublobatus Darlington, 1962
ssp. suffusus Darlington, 1962 [endemic]tor Darlington, 1971 [endemic]
truncatus (Nietner, 1858)Tachyta Kirby, 1837acuticollis Putzeys, 1875
coracina Putzeys, 1875
Limnastis Motschulsky, 1862atricapillus Bates, 1892
pilosus Bates, 1892
Tribus TrechiniPerileptus Schaum, 1860
japonicus Bates, 1873Perileptodes Jeannel, 1926darlingtoni Casale & Laneyrie, 1989
pilifer Jeannel, 1926
Tribus PsydriniMecyclothorax Sharp, 1903
bilaianus Baehr, 1998 [endemic]
eipomeki Baehr, 1995 [endemic]
eliti Baehr, 1995 [endemic]
jiwikae Baehr, 1995 [endemic]
riedeli Baehr, 1992 [endemic]
sapei Baehr, 1995 [endemic]
toxopei Darlington, 1962 [endemic]Tribe Pterostichini
Abacetus Dejean, 1828
haplosternus Chaudoir, 1878
convexiusculus Chaudoir, 1869
Analoma Darlington, 1971fortis (Darlington, 1962) [endemic]
gracilis (Darlington, 1962) [endemic]
Brachidius Chaudoir, 1852crassicornis Chaudoir, 1852Caelostomus Macleay, 1825subsinuatus (Chaudoir, 1833)Catadromus Macleay, 1825tenebroides (Olivier, 1790)Haploferonia Darlington, 1962simplex Darlington, 1962 [endemic]Homalonesiota Maindron, 1908Lesticus Dejean, 1828depressus Darlington, 1962 [endemic]
gracilis Darlington, 1962 [endemic]
medius Darlington, 1971 [endemic]
nitescens Sloane, 1907
politus Chaudoir, 1868
toxopei Darlington, 1962 [endemic]
Loxandrus Leconte, 1852major Darlington, 1962
medius Darlington, 1962
Morion Latreille, 1810humeratum Chaudoir, 1880
longipenne Putzeys, 1875
Nebrioferonia Straneo, 1939strigitarsis Straneo, 1939Platycoelus Blanchard, 1853archboldi Darlington, 1962
depressus Blanchard, 1853
Prosopogmus Chaudoir, 1865garivagliae Straneo, 1938Rhytiferonia Darlington, 1962iebele Darlington, 1962 [endemic]
julianae Baehr, 2001 [endemic]
nigra Darlington, 1962 [endemic]
oblongicollis Baehr, 2001 [endemic]
ophthalmica Baehr, 2001 [endemic]
paucistriata Baehr, 2001 [endemic]
punctigera Baehr, 2001 [endemic]
Stegazopteryx Will, 2004
ivimkaensis Will, 2004Tiferonia Darlington, 1962parva Darlington, 1962
Tribus PlatyniniAltagonum Darlington, 1952
erugatum Darlington, 1971 [endemic]
europhilum Darlington, 1952
grossulum Darlington, 1952
ssp. grossulum Darlington, 1952
ssp. intensum Darlington, 1952 [endemic]
ssp. reticolle Darlington, 1952 [endemic]noctellum Darlington, 1952
nox Darlington, 1952 [endemic]
regiscapha Darlington, 1952 [endemic]
scapha Darlington, 1952
sororium Darlington, 1971
sphodrum Darlington, 1952
stellaris Darlington, 1971 [endemic]
tutum Darlington, 1952 [endemic]
Arhytinus Bates, 1889
frater Baehr, 2010 [endemic]
medius Darlington, 1952
Cinctagonum Baehr, 2012
cordicolle Baehr, 2012 [endemic]Collagonum Baehr, 1995convexum Baehr, 1995 [endemic]
laticolle (Baehr, 1992)
ssp. laticolle (Baehr, 1992) [endemic]
ssp. macrops Baehr, 1995 [endemic]ophthalmicum (Baehr, 1992) [endemic]
riedeli Baehr, 1995 [endemic]
robustum Baehr, 1995 [endemic]Colpodes Macleay, 1825simplicauda Darlington, 1952
sinuicauda Darlington, 1952
Euplynes Schmidt-Goebel, 1846apicalis Darlington, 1952
laetus Darlington, 1952
Fortagonum Darlington, 1952acuticolle Baehr, 1995 [endemic]
angusticolle Baehr, 2010 [endemic]
bisetosiceps Baehr, 1995 [endemic]
bufo Darlington, 1952 [endemic]
curtum Baehr, 1992 [endemic]
cychriceps Darlington, 1952 [endemic]
denticulatum Baehr, 1995 [endemic]
depressum Baehr, 1995 [endemic]
forceps Darlington, 1952 [endemic]
formiceps Darlington, 1971 [endemic]
globulipenne Baehr, 1998 [endemic]
insulare Baehr, 2001 [endemic]
laevigatum Baehr, 1998 [endemic]
latum Baehr, 1995 [endemic]
sinak Baehr, 1998 [endemic]
skalei Baehr, 2008 [endemic]
spinipenne Baehr, 1998 [endemic]
spinosum Baehr, 1995 [endemic]
subconicolle (Darlington, 1971) [endemic]
sulcipenne Baehr, 2012 [endemic]
unipunctatum Baehr, 1995 [endemic]
Gastragonum Darlington, 1952esulcatum Baehr, 2012 [endemic]
laevisculptum Darlington, 1952 [endemic]
subrotundum Darlington, 1952
terrestroides Darlington, 1952 [endemic]
trechoides Darlington, 1952 [endemic]
Idiagonum Darlington, 1952capellae Baehr, 2012 [endemic]
darlingtoni Baehr, 2000 [endemic]
eliti Baehr, 2000 [endemic]
inasperum Darlington, 1952 [endemic]
latior Baehr, 2000 [endemic]
longipenne Baehr, 2000 [endemic]
muscorum Darlington, 1952 [endemic]
opacicolle Baehr, 2000 [endemic]
riedeli Baehr, 2000 [endemic]
sinuatipenne Baehr, 2000 [endemic]Iridagonum Darlington, 1952fessum Darlington, 1971
quadripunctellum Darlington, 1952
vigil Darlington, 1971 [endemic]Lithagonum Darlington, 1952Lorostemma Motschulsky, 1864informalis Darlington, 1952Maculagonum Darlington, 1952altipox Darlington, 1952
ssp. pallipox Darlington, 1952 [endemic]depilapox Darlington, 1971
plagipox Darlington, 1952
pox Darlington, 1952 [endemic]
seripox Darlington, 1971Montagonum Darlington, 1952riedeli Baehr, 2008 [endemic]
toxopeanum Darlington, 1952 [endemic]
Nebriagonum Darlington, 1952persetosum Baehr, 2012 [endemic]
subcephalum Darlington, 1952 [endemic]
Notagonum Darlington, 1952addendum Darlington, 1952
aitape Darlington, 1952
ssp. aitape Darlington, 1952
ssp. sansapor Darlington, 1952 [endemic]angustellum Darlington, 1952
astrum Darlington, 1971 [endemic]
crenulipenne Baehr, 2010 [endemic]
darlingtoni Baehr, 2010 [endemic]
dentellum Darlington, 1952
devosi Baehr, 2010 [endemic]
excisipenne Baehr, 2010
externum Darlington, 1952
fuscipes Baehr, 2010 [endemic]
gibbum Darlington, 1952 [endemic]
hamatum Baehr, 2010 [endemic]
ilagae Baehr, 2010 [endemic]
iridior Darlington, 1952 [endemic]
kitchingi Baehr, 2010
lackneri Baehr, 2010 [endemic]
laticolle Baehr, 2010 [endemic]
macrophthalmum Baehr, 2010
malkini Darlington, 1952 [endemic]
margaritum Darlington, 1952
ssp. margaritum Darlington, 1952
ssp. digulense Baehr, 2010 [endemic]
ssp. planipenne Baehr, 2010 [endemic]
ssp. montorum Baehr, 2010 [endemic]marginale Baehr, 2010 [endemic]
nigrinum Baehr, 2010 [endemic]
pernitidum Baehr, 2008 [endemic]
quadruum Darlington, 1971
reversior Darlington, 1952 [endemic]
schuelei Baehr, 2010 [endemic]
sigi Darlington, 1952 [endemic]
sinuum Darlington, 1952
subangulum Darlington, 1952 [endemic]
subimpressum Darlington, 1952
subpunctum Darlington, 1952
ssp. subpunctum Darlington, 1952
ssp. capitis Darlington, 1952 [endemic]subrufum Darlington, 1952
vaporum Darlington, 1952 [endemic]
vile Darlington, 1952
weigeli Baehr, 2008 [endemic]
Plicagonum Darlington, 1952fulvum Darlington, 1952 [endemic]
rugifrons Darlington, 1952
Potamagonum Darlington, 1952julianae Darlington, 1971 [endemic]Sericoda Kirby, 1837
ceylanicum (Motschulsky, 1859)Violagonum Darlington, 1956
Tribus PerigoniniPerigona Castelnau, 1835
aterrima Baehr, 2013 [endemic]
brachypennis Baehr, 2013 [endemic]
darlingtoni Baehr, 2013
erimae Csiki, 1924
fakfak Baehr, 2013 [endemic]
glabripennis Baehr, 2013 [endemic]
kaimanae Baehr, 2013 [endemic]
litura (Perroud & Montrouzier, 1864)
ludovici Csiki, 1924
ssp. ludovici Csiki, 1924nigriceps (Dejean, 1831)
ssp. nigriceps (Dejean, 1831)obtusangula Baehr, 2013 [endemic]
paralivens Baehr, 2013
plagiata Putzeys, 1875
proxima Baehr, 2013
pygmaea Andrewes, 1930
rex Darlington, 1968
riedeli Baehr, 2013 [endemic]
rossi Darlington, 1968
rubricincta Baehr, 2013 [endemic]
rufocastanea Baehr , 2013 [endemic]
subcordata Putzeys, 1875
subcyanescens Putzeys, 1875
suturalis Putzeys, 1875
transgrediens Baehr, 2013
Tribus Harpalini
Acupalpus Latreille, 1829exactellus Darlington, 1968 [endemic]
exactus Darlington, 1968
papua Darlington, 1968
ustus Andrewes, 1930
Chydaeus Chaudoir, 1854papua Darlington, 1968Coleolissus Bates, 1892angulatus Darlington, 1968
papua Darlington, 1968
Egadroma Motschulsky, 1855cyclops Darlington, 1968 [endemic]
quinquepustulatum (Wiedemann, 1823)
robustum Sloane, 1907
smaradgulum (Fabricius, 1798)
Gnathaphanus Macleay, 1825licinoides Hope, 1842
upolensis (Csiki, 1915)
Harpaloxenus Schauberger,1933celebensis Schauberger, 1933
fortis Darlington, 1968
mas Darlington, 1968
sedlaceki Darlington, 1968
Hyphaereon Macleay, 1825levis Darlington, 1968 [endemic]
timidus Darlington, 1968
Lecanomerus Chaudoir, 1850angustior Darlington, 1968 [endemic]Lyter Darlington, 1968glaber Darlington, 1968Notiobia Perty, 1830papuella (Darlington, 1968)
papuensis (Macleay, 1976)
Platymetopus Dejean, 1829Stenolophus Stephens, 1828volucer Andrewes, 1930Trichotichnus Morawitz, 1863delicatus Darlington, 1968
denarius Darlington, 1968
Tribus LicininiBadister Clairville, 1806
sundacicus Andrewes, 1926Dicrochile Guérin, 1846acuta Darlington, 1968
alternans Darlington, 1968
tiro Darlington, 1968 [endemic]Microferonia Blackburn, 1890alticola Baehr, 1998 [endemic]
avicapitis Baehr, 1998 [endemic]
habbemae Baehr, 1998 [endemic]Omestes Andrewes, 1933torta Andrewes, 1933Physoloesthus Chaudoir, 1850
caviceps (Andrewes, 1936)
Tribus Anaulacini
Aephnidius Macleay, 1825
adeloides Macleay, 1825Anaulacus Macleay, 1825
siamensis Chaudoir, 1876Odontomasoreus Darlington, 1968
humeralis Darlington, 1968
ssp. reductus Darlington, 1968
Tribus Chlaeniini
Chlaenius Bonelli, 1810
bimaculatus Dejean, 1826
ssp. pongraczi Jedlicka, 1951daer Darlington, 1968
hamifer Chaudoir, 1856
ssp. malcheri Van Emden, 1937maculiger Castelnau, 1867
occultus Sloane, 1907
olthofi Darlington, 1968
pan Darlington, 1968
siccus Darlington, 1968
Hololeius Laferté, 1851ceylanicus (Nietner, 1856)
Tribus Pentagonicini
Pentagonica Schmidt-Göbel, 1846
angulipennis Baehr, 2012 [endemic]
apiceincisa Baehr, 2012
avicapitis Baehr, 2012 [endemic]
batantae Baehr, 2012 [endemic]
biak Baehr, 2012 [endemic]
cognata Baehr, 2012
communis Baehr, 2012
convexipennis Baehr, 2012 [endemic]
cyanea (Montrouzier, 1860)
darlingtoni Baehr, 2012 [endemic]
erichsoni Schmidt-Goebel, 1846
estriata Darlington, 1968
fakfak Baehr, 2012 [endemic]
gressitti Baehr, 2012 [endemic]
lucens Baehr, 2012 [endemic]
macrops Baehr, 2012 [endemic]
novaeguineae Baehr, 2012
obscuripes Baehr, 2012 [endemic]
orbitalis Baehr, 2012 [endemic]
pallipes (Nietner, 1856)
profemorata Baehr, 2012
riedeli Baehr, 2012 [endemic]
semilaevis Baehr, 2012
similis Baehr, 2012
weigeli Baehr, 2012
Scopodes Erichson, 1842altus Darlington, 1968 [endemic]
amplipennis Baehr, 1995 [endemic]
arfakensis Baehr, 2010 [endemic]
balkei Baehr, 1995 [endemic]
bicolor Baehr, 1994 [endemic]caeruleus Baehr, 1994 [endemic]
chalceus Baehr, 1994 [endemic]cheesmannae Darlington, 1968 [endemic]
cuprascens Baehr, 1994 [endemic]foveipennis Baehr, 1994
interruptus Baehr, 1998 [endemic]laevifrons Baehr, 1994 [endemic]
minor Baehr, 1994muliae Baehr, 1995 [endemic]
perignitus Baehr, 1998 [endemic]reticulatus Baehr, 1994 [endemic]
riedeli Baehr, 1994 [endemic]rufipes Baehr, 1994 [endemic]striaticollis Baehr, 1994 [endemic]
tristis Baehr, 1994 [endemic]violaceus Baehr, 1994 [endemic]
Tribus Hexagoniini
Hexagonia Kirby, 1825
gressitti Darlington, 1971 [endemic]
Tribus OdacanthiniClarencia Sloane, 1917
papua Darlington, 1968
quadridens Darlington, 1968
Crassacantha Baehr, 1995
bidens Baehr, 1995 [endemic]Dicraspeda Chaudoir, 1862
bispinosa Darlington, 1968
denticulata Baehr, 1997
dubia (Gestro, 1879)
glabripennis Baehr, 2006 [endemic]
intermedia Baehr, 1997 [endemic]
laticollis Baehr, 1997 [endemic]
longiloba (Liebke, 1938)
obsoleta Baehr, 1996 [endemic]
ophthalmica Baehr, 2009 [endemic]
quadrispinosa Chaudoir, 1869
ssp. quadrispinosa Chaudoir, 1869Dobodura Darlington, 1968
armata Darlington, 1968Polydamasium Liebke, 1938
anomalum (Darlington, 1968)
nitidipennis Baehr, 2011 [endemic]Colasidia Basilewsky, 1954
avicapitis Baehr, 2011 [endemic]Tribus Helluonini
Helluonidius Chaudoir, 1872
chrysocomes Maindron, 1908
latipes Darlington, 1968
politus Darlington, 1968
Helluopapa Darlington, 1968
cheesmani Darlington, 1971 [endemic]
toxopei Darlington, 1968 [endemic]
Helluodema Castelnau, 1867
Tribus PhysocrotaphiniPogonoglossus Chaudoir, 1862
glabricollis Van Emden, 1937
laevissimus Baehr, 1995 [endemic]
latior Darlington, 1968 [endemic]
minor Darlington, 1968
papua Darlington, 1968
parvus Darlington, 1968
taylori Darlington, 1968
Schuelea Baehr, 2004
arfakensis (Baehr, 1987) [endemic]
monstrosa Baehr, 2004 [endemic]
Tribus LebiiniAnomotarus Chaudoir, 1875
Aristolebia Bates, 1892
capitis Darlington, 1968 [endemic]
mucronata (Sloane, 1907)
Catascopus Kirby, 1825
aruensis Saunders, 1863
astrum Baehr, 2012 [endemic]
biroi Darlington, 1968
brunneus Darlington, 1968
dalbertisi Straneo, 1943
elegans (Weber, 1801)
ssp. amoenus Chaudoir, 1861
ssp. biakensis Baehr, 2012 [endemic]latus Darlington, 1968 [endemic]
moorei Straneo, 1994
platypennis Baehr, 2012
riedeli Baehr, 1997 [endemic]
sidus Darlington, 1968
ssp. darlingtoni Baehr, 2012 [endemic]
ssp. japensis Baehr, 2012 [endemic]strigicollis Straneo, 1943
waigeoensis Baehr, 2012 [endemic]
weigeli Baehr, 2012
Celaenephes Schmidt-Göbeöl, 1846
linearis (Walker, 1858)
parallelus Schmidt-Göbel, 1846
Cheilagona Baehr, 2006
gressitti (Darlington, 1968)
ssp. gressitti (Darlington, 1968)
ssp. planata Baehr, 2006 [endemic]nigropicea Baehr, 2006 [endemic]
variabilis (Darlington,1968)
Coptodera Dejean, 1825
cyanella Bates, 1869
eluta Andrewes, 1923
lineolata Bates, 1869
ornatipennis Louwerens, 1962
oxyptera Chaudoir, 1869
papuella Darlington, 1968
Demetrida White, 1846
aitape Darlington, 1968
basalis Darlington, 1968 [endemic]
brunnea Darlington, 1968
discoidalis Darlington, 1968
divisa Darlington, 1968
duplicata Darlington, 1968
forma Darlington, 1968
genicula Darlington, 1968
hollandia Darlington, 1968
humeralis Darlington, 1968
imitatrix Darlington, 1968
kokoda Darlington, 1968
latangula Darlington, 1968
magna Darlington, 1968
nigriceps Darlington, 1968
nigripennis Darlington, 1968 [endemic]
nigripes Darlington, 1968
nubicola Darlington, 1968 [endemic]
pallens Darlington, 1968 [endemic]
parena Darlington, 1968 [endemic]
recta Darlington, 1968
reversa Darlington, 1968 [endemic]
saidor Darlington, 1968
sibil Darlington, 1968
sublepida Darlington, 1968
submoda Darlington, 1968
subtenuis Darlington, 1968
tesselata Darlington, 1968
truncata Darlington, 1968
viridibasis Darlington, 1968
viridipennis Darlington, 1968
wau Darlington, 1968
Dolichoctis Schmidt-Goebel, 1846
aculeata Chaudoir, 1869
angustemaculata Baehr, 1999
ssp. angustemaculata Baehr, 1999 [endemic]avicapitis Baehr, 1999 [endemic]
biak Baehr, 1999 [endemic]
bicolor Baehr, 1999 [endemic]
bisetosa Baehr, 1999
darlingtoni Baehr, 1999 [endemic]
divisa Darlington, 1968
exigua Baehr, 1999
glabripennis Baehr, 2003
lackneri Baehr, 2013 [endemic]
longicornis Baehr, 1999
major Baehr, 1999
nigricauda Baehr, 2007 [endemic]
ophthalmica Baehr, 1999 [endemic]
riedeli Baehr, 1999 [endemic]
spinosa Darlington, 1968
ssp. spinosa Darlington, 1968
ssp. longispina Baehr, 2006
ssp. subspina Baehr, 2006subquadrata Darlington, 1968
subrotunda Darlington, 1968
sulcicollis Baehr, 1999 [endemic]
suturalis Darlington, 1968
tetrastigma Chaudoir, 1869
ssp. papuana Baehr, 2013vixstriata Baehr, 1999 [endemic]
Holcoderus Chaudoir, 1869
Lebia Latreille, 1802
Microlestes Schmidt-Göbel, 1846
curtatus Darlington, 1968Minuphloeus Darlington, 1968
Minuthodes Andrewes, 1941
Miscelus Klug, 1834
sibling Darlington, 1968
unicolor Putzeys, 1875
Mochtherus Schmidt-Göbel, 1846
Oxyodontus Chaudoir, 1869
tripunctatus Chaudoir, 1869Parena Motschulsky, 1859
fasciata (Chaudoir, 1872)
picea (Macleay, 1871)
Pericalus Macleay, 1825
cuprascens Baehr, 1994 [endemic]
figuratus Chaudoir, 1861
Phloeocarabus Macleay, 1871
nigricollis (Macleay, 1864)Physodera Eschscholtz, 1829
cyanipennis v.d.Poll, 1889Pseudoplatiella Baehr, 2012
Sinurus Chaudoir, 1869
opacus Chaudoir, 1869Stenotelus Bouchard, 1903
Stricklandiana Bousquet, 2003
Syntomus Hope, 1834
quadripunctatus (Schmidt-Göbel, 1846)Trigonothops Macleay, 1864
lateralis Darlington, 1968 [endemic]
Tribus Pseudomorphini
Sphallomorpha Westwood, 1837cheesmannae Baehr, 1992 [endemic]
litterata Baehr, 1992
oculata Baehr, 2004 [endemic]
setifera Baehr, 1993
Tribe Oodini
Anatrichis Leconte, 1853
pusilla Sloane, 1910Oodes Bonelli, 1810cribristernis Bates, 1892
denisonensis Castelnau, 1867
exiguus Andrewes, 1933
longior Darlington, 1968
nil Darlington, 1968
par Darlington, 1968
piceus Nietner, 1856
rossi Darlington, 1968 [endemic]
siamensis Chaudoir, 1882
ssp. vulsus Darlington, 1968siccus Darlington, 1968
Tribus Cyclosomini
Sarothrocrepis Chaudoir, 1876
papua Darlington, 1968
Tribus Brachinini
Brachinus Weber, 1801
papua Darlington, 1968 [endemic]Pheropsophus Solier, 1833adrianae Giachino, 2005 [endemic]
aptinomorphus Heller, 1910
baehri Giachino, 2005 [endemic]
baliothorax Heller, 1910
balkei Giachino, 2005 [endemic]
brussinoi Giachino, 2005 [endemic]
pedes Darlington, 1968 [endemic]
riedeli Giachino, 2005
verticalis Dejean, 1825
wolfi Giachino, 2005 [endemic]
Checklist last updated on 10th February 2014
Literature
Baehr, M. 1987a. The Indo-Australian species of the Zuphiine genus Agastus Schmidt-Göbel (Insecta, Coleoptera, Carabidae). - Spixiana 10: 1-11.
Baehr, M. 1987b. Pogonoglossus arfakensis sp. nov. from New Guinea (Coleoptera, Carabidae, Helluodinae). - Dt. ent. Z., N. F. 34: 363-366.
Baehr, M. 1991. On new and rare Leleupidiini from the Oriental and Australian Regions (Coleoptera, Carabidae, Zuphiinae). - Mitt. Münch. Ent. Ges. 81: 193-202.
Baehr, M. 1992a. A new Mecyclothorax Sharp from New Guinea (Insecta, Coleoptera, Carabidae, Psydrinae). - Spixiana 15: 249-252.
Baehr, M. 1992b. On some agonine beetles of the genus Fortagonum DARLINGTON from New Guinea (Coleoptera, Carabidae, Agoninae). - Mitt. Münch. Ent. Ges. 82: 73-81.
Baehr, M. 1992c. Revision of the Pseudomorphinae of the Australian Region 1. The previous genera Sphallomorpha Westwood and Silphomorpha Westwood. Taxonomy, phylogeny, zoogeography (Insecta, Coleoptera, Carabidae). - Spixiana Suppl. 18: 1-440.
Baehr, M. 1993. A remarkable new species of the genus Sphallomorpha Westwood from Irian Jaya (New Guinea). 2. Supplement to the "Revision of the Pseudomorphinae of the Australian Region 1." (Insecta, Coleoptera, Carabidae). - Spixiana 16: 207-211.
Baehr, M. 1994a. Revision of the genus Scopodes Erichson from New Guinea (Insecta, Coleoptera, Carabidae, Pentagonicinae). - Spixiana 17: 97-155.
Baehr, M. 1994b. New species and new records of the genus Sphallomorpha Westwood from Australia and New Guinea. 3rd Supplement to the "Revision of the Pseudomorphinae of the Australian Region 1." (Insecta, Coleoptera, Carabidae). - Spixiana 17: 215-235.
Baehr, M. 1994c. Revision of the subgenus Coeloprosopus CHAUDOIR of the ground beetle genus Pericalus MACLEAY. Taxonomy, phylogeny, zoogeography (Coleoptera, Carabidae, Lebiinae). - Mitt. Münch. ent. Ges. 84: 13-58.
Baehr, M. 1995a. New species and new records of the genera Fortagonum Darlington and Collagonum, gen. nov. from New Guinea (Insecta, Coleoptera, Carabidae, Agoninae). - Spixiana 18: 15-43.
Baehr, M. 1995b. A new genus of Odacanthinae from New Guinea (Insecta, Coleoptera, Carabidae). - Spixiana 18: 45-48.
Baehr, M. 1995c. New taxa and new records of the genus Scopodes Erichson from New Guinea. Supplement to the "Revision of the genus Scopodes Erichson from New Guinea" (Insecta, Coleoptera, Carabidae, Pentagonicinae). - Spixiana 18: 111-121.
Baehr, M. 1995d. A peculiar new species of the genus Pogonoglossus Chaudoir from New Guinea (Insecta, Coleoptera, Carabidae, Helluodinae). - Spixiana 18: 255-258.
Baehr, M. 1995e. The genus Mecyclothorax Sharp in New Guinea (Insecta, Coleoptera, Carabidae, Psydrinae). - Mitt. Münch. ent. Ges. 85: 3-19.
Baehr, M. 1996a. A new species of the genus Anomotarus Chaudoir from New Guinea (Insecta, Coleoptera, Carabidae, Lebiinae). - Spixiana 19: 17-20.
Baehr, M. 1996b. Three new species of the genus Dicraspeda Chaudoir from New Guinea (Insecta, Coleoptera, Carabidae, Odacanthinae). - Spixiana 19: 137-146.
Baehr, M. 1996c. The ground beetle genus Casnoidea Castelnau. Taxonomy, phylogeny, zoogeography (Insecta, Coleoptera, Carabidae, Odacanthinae). - Invertebr. Taxon. 10: 1041-1084.
Baehr, M. 1996d. A peculiar new species of Anomotarus (Nototarus) from New Guinea (Coleoptera, Carabidae, Lebiinae). - Rev. Suisse Zool. 103: 607-611
Baehr, M. 1997a. Revision of the Pseudomorphinae of the Australian Region 2. The genera Adelotopus Hope, Cainogenion Notman, Paussotropus Waterhouse, Pseudomorpha Kirby, and Cryptocephalomorpha Ritsema. Taxonomy, phylogeny, zoogeography (Insecta, Coleoptera, Carabidae). - Spixiana Suppl. 23: 1-510.
Baehr, M. 1997b. The genus Stricklandia Macleay in New Guinea (Carabidae, Lebiinae). - Coleoptera 1: 117-136.
Baehr, M. 1997c. Review of the Australian-Papuan laevigatus-group of the genus Catascopus Kirby, with description of a new species (Insecta, Coleoptera, Carabidae, Lebiinae).- Spixiana 20: 227-234.
Baehr, M. 1997d. Three further new species of the genus Dicraspeda CHAUDOIR from New Guinea (Coleoptera, Carabidae, Odacanthinae). - Mitt. Münch. Ent. Ges. 87: 29-37
Baehr, M. 1998a. New species of the genus Fortagonum Darlington from western New Guinea (Insecta, Coleoptera, Carabidae, Agoninae). - Spixiana 21: 11-20.
Baehr, M. 1998b. A further new species of the genus Mecyclothorax Sharp from western New Guinea (Insecta, Coleoptera, Carabidae, Psydrinae). - Spixiana 21: 21-24.
Baehr, M. 1998c. New species and new records of the genus Scopodes Erichson from New Guinea (Insecta, Coleoptera, Carabidae, Pentagonicinae). - Spixiana 21: 145-158.
Baehr, M. 1998d. Two further new species of the genus Dicraspeda Chaudoir from New Guinea and the New Hebrides (Insecta, Coleoptera, Carabidae, Odacanthinae). - Entomofauna 19: 173-184.
Baehr, M. 1998e. Two new species of Minuthodes Andrewes from New Guinea (Insecta, Coleoptera, Carabidae, Lebiinae). - Spixiana 21: 235-240.
Baehr, M. 1998f. The genus Microferonia Blackburn in New Guinea (Insecta, Coleoptera, Carabidae, Licininae). - Mitt. Münch. ent. Ges. 88: 25-32.
Baehr, M. 1999a. New species and new records of the genus Scopodes Erichson from New Guinea. Third Supplement to the "Revision of the genus Scopodes Erichson from New Guinea". (Insecta, Coleoptera, Carabidae, Pentagonicinae). - Spixiana 22: 97-101.
Baehr, M. 1999b. The genus Dolichoctis Schmidt-Göbel in New Guinea and surrounding areas (Insecta, Coleoptera, Carabidae, Lebiinae). - Coleoptera 2: 121-169 (1998).
Baehr, M. 2000a. Revision of the genus Idiagonum Darlington from New Guinea (Carabidae, Agoninae). - Coleoptera 4: 55-92.
Baehr, M. 2000b. A new species of the leleupidiine genus Colasidia Basilewsky from New Guinea (Coleoptera, Carabidae, Zuphiinae). - Spixana 23: 41-45.
Baehr, M. 2000c. Some genera and species of ground beetles new to Australia (Coleoptera: Carabidae). - Mem. Qld. Mus. 46: 9-14.
Baehr, M. 2001a. Further new species of the genera Fortagonum Darlington and Collagonum Baehr from New Guinea (Insecta, Coleoptera, Carabidae, Agoninae) – Spixiana 24: 53-72.
Baehr, M. 2001b. The genus Rhytiferonia Darlington in New Guinea (Carabidae: Pterostichinae). – Coleoptera 5: 41-55.
Baehr, M. 2002a. Herculagonum atlas, gen. et spec. nov. from Papua New Guinea (Insecta, Carabidae, Platyninae). – Spixiana 25: 131-135
Baehr, M. 2002b. Two new species of the genus Mecyclothorax Sharp from Papua New Guinea (Insecta, Coleoptera, Carabidae, Psydrinae). – Rev. Suisse Ent. 109: 695-704.
Baehr, M. 2003a. Further new species of the genus Dolichoctis Schmidt-Göbel from New Guinea and surrounding islands (Insecta, Coleoptera, Carabidae, Lebiinae). - Anim. Biodiv. Cons. 26: 1-7.
Baehr, M. 2003b. A revision of the brunnea-group of the genus Dicraspeda Chaudoir (Coleoptera, Carabidae, Odacanthinae). – Spixiana 26: 249-267.
Baehr, M. 2003c. New records of the genus Dolichoctis Schmidt-Göbel from New Guinea and surrounding islands (Insecta, Coleoptera, Carabidae, Lebiinae). – Arxius Miscel. Zool.1: 12-17.
Baehr, M. 2004a. A new genus and two new species of Physocrotaphinae from New Guinea (Insecta, Coleoptera, Carabidae). – Spixiana 27. 107-113.
Baehr, M. 2004b. A peculiar new species of the genus Sphallomorpha WESTWOOD from New Guinea (Coleoptera: Carabidae: Pseudomorphinae). – Koleopt. Rundschau 74: 25-31.
Baehr, M. 2004c. The genus Lebia Latreille in the Australian Region (Insecta, Coleoptera, Carabidae, Lebiinae). – Spixiana 27: 205-246.
Baehr, M. 2004d. A new species of the genus Aristolebia Bates from Thailand, with notes on some Papuan and Australian species (Insecta, Coleoptera, Carabidae, Lebiinae). – Spixiana 27: 247-251.
Baehr, M. 2004e. A revision of the Oriental, New Guinean, and Pacific species of the ground beetle genus Anomotarus Chaudoir (Coleoptera: Carabidae: Lebiinae) – Coleoptera 7: 27-78 (2003).
Baehr, M. 2004f. A revision of the Australian odacanthine ground beetles, including Checklists for Australia and the Papuan Subregion (Insecta: Coleoptera: Carabidae). – Mem. Qld. Mus. 50: 133-194.
Baehr, M. 2004g. Colasidia wau, a new leleupidiine species from Papua New Guinea (Insecta, Coleoptera, Carabidae, Zuphiinae). – Rev. Suisse Zool. 111: 175-181.
Baehr, M. 2005a. New species and new records of the genus Anomotarus Chaudoir, subgenus Anomotarus s. str. from New Guinea (Insecta, Coleoptera, Carabidae, Lebiinae). – Spixiana 28: 169-173.
Baehr, M. 2005b. Two new species of the genus Pogonoglossus Chaudoir from New Guinea (Coleoptera, Carabidae, Physocrotaphinae). – Bull. Inst. Roy. Sci. Nat. Belg. 75: 5-12.
Baehr, M. 2006a. New species and new records of the genera Dicraspeda Chaudoir and Eudalia Castelnau from the Papuan and Australian regions, with a nomenclatorial note on Deipyrus Liebke (Insecta, Coleoptera, Carabidae, Odacanthinae). – Spixiana 29: 51-72.
Baehr, M. 2006b. Revision of the genera Agonocheila Chaudoir and Minuthodes Andrewes in New Guinea (Insecta, Coleoptera, Carabidae, Lebiinae). – Spixiana 29: 103-145.
Baehr, M. 2006c. New taxa of the lebiine genus Dolichoctis and of a related new genus, from the Papuan subregion (Coleoptera: Carabidae: Lebiinae). – Folia Heyrovskyana 14: 37-66.
Baehr, M., 2008a. Two new species of the genus Mecyclothorax Sharp from New Guinea (Coleoptera: Carabidae: Psydrinae). Tijdschrift voor Entomologie 151: 133-140.
Baehr, M., 2008b. New agonine species from New Guinea (Coleoptera: Carabidae: Agonini). – Folia Heyrovskyana 16: 55-76.
Baehr, M. 2008c. The Australian Clivinini 1. The genera Ancus Putzeys, Aspidoglossa Putzeys, Clivinarchus Sloane, Platysphyrus Sloane, Pseudoclivina Kult, Rhysocara Sloane, Syleter Andrewes, the subgenera Paraclivina Kult, Semiclivina Kult, and the atrata-, biplagiata-, brevicornis-, coronata-, coryzoides-, cribrosa-, denticollis-, grandiceps-, incerta-, lobata-, obliquata-, obsoleta, orbitalis-, planiceps-, sulcaticeps-, tranquebaria-, and wurargae-groups of the genus Clivina Latreille. With a note on a record of the genus Parathlibops Basilewsky (Scapterini) (Carabidae, Scaritinae). – Coleoptera 12: 1-220.
Baehr, M. 2009a. A new species of the genus Adelotopus Hope from the Moluccas. With new records of some New Guinean species of the genus Sphallomorpha Westwood. 10th Supplement to the "Revision of the Pseudomorphinae of the Australian Region." (Insecta, Coleoptera, Carabidae, Pseudomorphinae). – Spixiana 32: 39-43.
Baehr, M. 2009b. Further taxonomic differentiation in the quadrispinosa complex of the genus Dicraspeda Chaudoir in the Papuan Region; with new records of other species of Dicraspeda (Coleoptera: Carabidae: Odacanthinae) – Tijdschrift voor Entomologie 152: 99-108.
Baehr, M. 2009c. New genera and species and new records of odacanthine carabid beetles from the Australian, Papuan, and Oriental Regions (Insecta, Coleoptera, Carabidae, Odacanthini). – Spixiana 32: 201-229.
Baehr, M., 2010a. A new species of the genus Fortagonum Darlington from New Guinea (Coleoptera: Carabidae: Platynini). Tijdschrift voor Entomologie 153: 79-83.
Baehr, M. 2010b. A new species of the genus Scopodes Erichson from western New Guinea (Insecta, Coleoptera, Carabidae, Pentagonicini). Fourth Supplement to the "Revision of the genus Scopodes Erichson from New Guinea". – Spixiana 33: 55-58.
Baehr, M. 2010c. New species of the genus Notagonum Darlington from New Guinea related to N. angustellum Darlington (Insecta, Coleoptera, Carabidae, Platyninae). – Spixiana 33: 195-228.
Baehr, M. 2010d. New taxa of the genus Notagonum Darlington from New Guinea related to N. margaritum Darlington (Coleoptera, Carabidae, Platynini). – Spixiana 33: 229-245.
Baehr, M. 2010e. A revision of the genus Arhytinus BATES (Coleoptera: Carabidae: Platynini). – Bulletin de l’Institute Royale des Sciences Naturelles de Belgique 80: 5-50
Baehr, M. 2010f. Revision of the genus Scopodes Erichson in Australia (Insecta, Coleoptera, Carabidae, Pentagonicinae). – Coleoptera 14: 1-188
Baehr, M. 2010g. Two new species of the genus Lioptera Chaudoir from the Papuan and southern Oriental Regions (Coleoptera: Carabidae: Lebiinae). – Australian Entomologist 36: 171-182.
Baehr, M. 2011a. New species and new records of the genus Anomotarus Chaudoir, subgenus Anomotarus s. str., from the Oriental and Australian Regions. Supplement to the “Revisions of the species of the subgenus Anomotarus Chaudoir s. str.” (Insecta, Coleoptera, Carabidae, Lebiinae). – Spixiana 34: 21-32.
Baehr, M. 2011b. A new species of the genus Stricklandiana Bousquet (formerly Stricklandia Macleay) from New Guinea (Coleoptera, Carabidae, Lebiinae). – Tijdschrift voor Entomologie 154: 167-171.
Baehr, M. 2011c. Two new species of the genus Agastus Schmidt-Goebel from New Guinea and the Moluccas (Insecta, Coleoptera, Carabidae, Zuphiinae). – Tijdschrift voor Entomologie 154: 291-297.
Baehr, M. 2011d. Two new species of the leleupidiine genus Colasidia BASILEWSKY from New Guinea (Coleoptera, Carabidae, Zuphiini). – Mitteilungen der Münchner Entomologischen Gesellschaft 101: 59-67.
Baehr, M. 2012a. The genus Pentagonica Schmidt-Goebel in the Oriental, Papuan, and Australian Regions (Insecta, Coleoptera, Carabidae, Pentagonicini). – Coleoptera 15: 1-200 (2011)
Baehr, M. 2012b. New genera and species of platynine carabid beetles from New Guinea (Insecta, Coleoptera, Carabidae, Platynini). – Spixiana 35: 35-77
Baehr, M. 2012c. Revision of some species-groups of the genus Catascopus Kirby from the Oriental and Australian Regions (Coleoptera, Carabidae, Lebiini). – Coleoptera 108: 11-108
Baehr, M. 2013a. The species of the genus Perigona Castelnau from New Guinea, Sulawesi, Halmahera, and Australia, and of the parvicollis-pygmaea-lineage. (Coleoptera, Carabidae, Perigonini). – Entomologische Blätter und Coleoptera 109: 1-132.
Baehr, M. 2013b. The Dolichoctis striata complex (Coleoptera, Carabidae, Lebiini). – Entomologische Blätter und Coleoptera 109: 133-222.
Baehr, M. 2013c. Supplement to “The species of the genus Perigona Castelnau from New Guinea, Sulawesi, Halmahera, and Australia, and of the parvicollis-pygmaea-lineage. (Coleoptera, Carabidae, Perigonini)” and to “The Dolichoctis striata complex (Coleoptera, Carabidae, Lebiini). – Entomologische Blätter und Coleoptera 109: 223-231.
Baehr, M. 2013d. New species of the genus Dolichoctis Schmidt-Goebel, subgenus Spinidolichoctis Baehr, from the Papuan and Oriental Regions (Insecta: Coleoptera: Carabidae: Lebiinae). – Annals of the Carnegie Museum 82: 149-162
Darlington, P. J. Jr. 1952. The carabid beetles of New Guinea. Part 2. The Agonini. - Bull. Mus. comp. Zool. 107: 89-252.
Darlington, P. J. Jr. 1962. The carabid beetles of New Guinea. Part I. Cicindelinae, Carabinae, Harpalinae through Pterostichini. - Bull. Mus. comp. Zool. 126: 321-565.
Darlington, P. J. Jr. 1968. The Carabid beetles of New Guinea. Part III. Harpalinae continued. Perigonini to Pseudomorphini. – Bull. Mus. Comp. Zool. 139: 1-253.
Darlington, P. J. Jr. 1971. The carabid beetles of New Guinea. Part IV. General Considerations; Analysis and History of Fauna; Taxonomic Supplement. – Bull. Mus. comp. Zool. 142: 129-337.
Giachino, P. M. 2005. The genus Pheropsophus Solier, 1833 in New Guinea (Insecta, Coleoptera, Carabidae, Brachininae). – Spixiana 28: 223-257.
Will, K. W. 2004. A new genus of Caelostomini (Coleoptera: Carabidae) from Papua New Guinea. – The Coleopterists Bulletin 58: 637-641